Hypsiboas riojanus

Hypsiboas riojanus (Koslowsky, 1895)


Class: Amphibia > Order: Anura > Family: Hylidae > Genus: Hypsiboas > Species: Hypsiboas riojanus


Synonym: Hypsiboas andinus — Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005

English names:



A moderate-sized frog with: (1) snout rounded in dorsal view and bluntly rounded in profile; (2) skin on dorsum smooth to granular; (3) width of disc on Finger III less than diameter of tympanum; (4) tarsal fold low; (5) dorsal color pattern extremely variable consisting of small irregular spots; (6) flanks extremely variable small dark spots, pale blotches, uniform; (7) posterior surfaces of thighs with dark vertical bars.

The color pattern is highly variable and seems to show some geographic variation, the differences in color can change in the same locality and also in the same individual depending of the light and temperature. Eg: Dorsum uniformly dark or pale green or brown, with or without dark and irregular blotches; labial, canthal, and supratympanic stripes bronze; dorsolateral area with creamy-with stripe; flanks and groin blue with black spots; thighs and shanks with black and blue transverse bars.

Type locality:



This species is widely distributed in the Andean regions of Bolivia and Argentina. In the northern part of its range it occurs at elevations of 1650-4500 m, but in the southern part, it occurs at elevations of 500-1640 m. At high elevations,

La Paz, Oruro, Potosi, Cochabamba, Tarija, Chuquisaca and Santa Cruz.

Ecoregions: Humid Puna, Semi Humid Puna, Interandean Dry Valleys, Prepuna, Mountain Cloud forest.



The frog inhabits montane grassland; in the northern part of its range it descends into cloud forest and temperate forest, whereas in the southern part of its range it occurs in premontane deciduous forest.



The tadpole of Hyla andina was described and illustrated as H, pulchella andina by Cei (1980) and Lavilla (1984).




Water quality and habitat:

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External links:







Barrio, A. (1965). Las subespecies de Hyla pulchella Dumeril y Bibron (Anura, Hylidae). Physis, 25, 115–128.


De la Riva, I., Kohler, J., Lotters, S. & Reichle, S. (2000). Ten years of research on Bolivian amphibians: updated checklist, distribution, taxonomic problems, literature and iconography. Revista Española de Herpetología, 14, 19–164.



Duellman, W. E., De la Riva, I. & Wild, E. R. (1997). Frogs of the Hyla armata and Hyla puchella groups in the Andes of South America, with definitions and analyses of phylogenetic relationship of Andean groups of Hyla. Scientific Papers Natural History Museum University of Kansas, 3, 1–41.


Faivovich, J., Garcia, P. C. A., Ananias, F., Lanari, L., Basso, N. G. & Wheeler, W. C. (2004). A molecular perspective on the phylogeny of the Hyla pulchella species group (Anura, Hylidae). Molecular Phylogenetics and Evolution, 32, 938–950.


Faivovich, J., Haddad, C. F. B., Garcia, P. C. A., Frost, D. R., Campbell, J. A. & Wheeler, W. C. (2005). Systematic review of the frog family Hylidae, with special reference to Hylinae: phylogenetic analysis and taxonomic revision. Bulletin of the

American Museum of Natural History, 294, 1–240


Kohler, J., John, A. & Bo¨hme, W. (2006). Notes on amphibians recently collected in the Yungas de La Paz region, Bolivia. Salamandra, 42, 21–27


Köhler, J., Koscinski, D., Padial, J. M., Chaparro, J. C., Handford, P., Lougheed S. C. & De la Riva, I. 2010. Systematics of Andean gladiator frogs of the Hypsiboas pulchellus species group (Anura, Hylidae). Zoologica Scripta 36: 572-590.



Koscinski, D., Handford, P., Tubaro, P. L., Sharp, S. & Lougheed, S. C. (2008). Pleistocene climatic cycling and diversification of the Andean treefrog, Hypsiboas andinus. Molecular Ecology, 17, 2012–2025.


Langone, J. A. & Lavilla, E. O. (2002). Comentarios nomenclatoriales

sobre algunos taxa del grupo de Hyla pulchella (Anura: Hylidae). Cuadernos de Herpetologı´a, 16, 73–78.